He also questioned why the IOC had not allowed Russian whistleblower Yulia Stepanova to run as an independent athlete in Rio.

"Ms. Stepanova was instrumental in courageously exposing the single biggest doping scandal of all time," Niggli said. "WADA is very concerned by the message that this sends whistleblowers for the future."

Evidence from the McLaren report had also been sent to the International Paralympic Committee (IPC), WADA said.

The IPC will make its own decision on whether to permit Russian athletes in the Paralympics. It has already started suspension proceedings against Russia following the McLaren report.

Responding to IOC calls that the agency review the anti-doping system, WADA said it was fully committed to further strengthening system.

It also confirmed professor Richard McLaren would continue his investigative work into doping and "identify athletes that might have benefited from manipulation of the doping control process to conceal positive doping tests".